Zachary Shaffer

Position Title

PhD Candidate

Joint Appointments

 MD/PhD Program

Areas of Interest

Host-Microbiome Interactions in Pregnancy: An Immunobiological Perspective



  1. Greenberg JM, Romero R, Winters AD, Galaz J, Garcia-Flores V, Arenas-Hernandez M, Panzer J, Shaffer Z, Kracht DJ, Gomez-Lopez N, Theis KR. Microbiota of the Pregnant Mouse: Characterization of the Bacterial Communities in the Oral Cavity, Lung, Intestine, and Vagina through Culture and DNA Sequencing. Microbiol Spectr. 2022 Aug 31;10(4):e0128622. doi: 10.1128/spectrum.01286-22. PMID: 35916526
  2. Gomez-Lopez N, Galaz J, Miller D, Farias-Jofre M, Liu Z, Arenas-Hernandez M, Garcia-Flores V, Shaffer Z, Greenberg JM, Theis KR, Romero R. The immunobiology of preterm labor and birth: intra-amniotic inflammation or breakdown of maternal-fetal homeostasis. Reproduction. 2022 Jun 20;164(2):R11-R45. doi: 10.1530/REP-22-0046. PMID: 35559791
  3. Farias-Jofre M, Romero R, Galaz J, Xu Y, Tao L, Demery-Poulos C, Arenas-Hernandez M, Bhatti G, Liu Z, Kawahara N, Kanninen T, Shaffer Z, Chaiworapongsa T, Theis KR, Tarca AL, Gomez-Lopez N. Pregnancy tailors endotoxin-induced monocyte and neutrophil responses in the maternal circulation. Inflamm Res. 2022 Apr 21:1-16. doi: 10.1007/s00011-022-01569-z. PMID: 35445873
  4. Winters AD, Romero R, Greenberg JM, Galaz J, Shaffer ZD, Garcia-Flores V, Kracht DJ, Gomez-Lopez N, Theis KR. Does the Amniotic Fluid of Mice Contain a Viable Microbiota? Front Immunol. 2022 Feb 28;13:820366. doi: 10.3389/fimmu.2022.820366. PMID: 35296083
  5. Damico ME, Rueppell O, Shaffer Z, Han B, Raymann K. High royal jelly production does not impact the gut microbiome of honey bees. Anim Microbiome. 2021 Sep 13;3(1):60. doi: 10.1186/s42523-021-00124-1. PMID: 34517918
  6. Raymann K, Coon KL, Shaffer Z, Salisbury S, Moran NA. Correction for Raymann et al., "Pathogenicity of Serratia marcescens Strains in Honey bees". mBio. 2019 Feb 5;10(1):e02855-18. doi: 10.1128/mBio.02855-18. PMID: 30723134
  7. Raymann K, Coon KL, Shaffer Z, Salisbury S, Moran NA. Pathogenicity of Serratia marcescens Strains in Honey Bees. mBio. 2018 Oct 9;9(5):e01649-18. doi: 10.1128/mBio.01649-18. PMID: 30301854
  8. Raymann K, Shaffer Z, Moran NA. Antibiotic exposure perturbs the gut microbiota and elevates mortality in honeybees. PLoS Biol. 2017 Mar 14;15(3):e2001861. doi: 10.1371/journal.pbio.2001861. eCollection 2017 Mar. PMID: 28291793





 2017 - BS/BS in Biochemistry and Biology, University of Texas Austin


Dr. Kevin Theis

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